quad.ru.ma.nousadj.Zoology (形容词)【动物学】 Having four feet with opposable first digits, as primates other than human beings.四足如手的:有四足和与其他脚趾相对的第一个足趾的;如除人类以外的灵长类动物来源:quadru- quadru-Latin manus [hand] * see man- 2拉丁语 manus [手] *参见 man- 2-ous -ous